Conus
(Lithoconus) hendersoni (Marwick, 1931)
Description (3):
Large for New Zealand Conus species (to about 70 mm high), with
very low, stepped, conic spire arid very long, tapered, straight-sided last
whorl, long, narrow, parallel-sided aperture with shallow U-shaped sinus on the
sutural ramp, and prominent but low, smooth, rounded carina around shoulder of
all whorls (i.e., descending spire and forming steps in outline). Apart from
growth lines, and few weak, variable, indistinct spiral grooves high on whorls,
sculpture is of about 10 low, wide spiral cords on base of last whorl,
increasing in prominence down shell. Protoconch not seen.
Comparison (3):
Conus hendersoni is the youngest large Conus species in New
Zealand Cenozoic rocks. It may be related to such relatively cool-water living
species as C. teramachii (Kuroda, 1956), recorded by Marshall (1981b, p.
499) living at present just to the north of northernmost New Zealand, in
357-677 m; C. teramachii has a similar stepped, carinate spire and basal
spiral cords to those of C. hendersoni, but the spire is taller and the
carina nodulous in C. teramachii. The presence of a single common
species of Conus in Late Miocene rocks near East Cape need only indicate
sea temperatures a little warmer than those in the region at present, and only
the rather more diverse (but largely undescribed) Conus faunas of
Clifden, Southland (Altonian-Clifdenian) and Kaipara Harbour, Northland
(Otaian-Altonian) indicate markedly warmer temperatures. Conus is an
epifaunal toxoglossan carnivore, feeding on polychaete worms, molluscs or (in a
few species) fish, "stinging" its prey with individual, unattached
radular teeth and a complex toxin from a well developed poison gland, and the very
poisonous fish-eating species of the modern tropics are well known to have
killed a few humans. Most species occur on hard substrates in shallow water (on
rocky shores and coral reefs), but C. hendersoni has been collected
only from soft-substrate environments in mid-outer shelf depths.
The phylogeny of Conus has been investigated by several recent
authors, all indicating an essentially simple taxonomy with only two genera.
Most recently, Bandyopadhyay et al. (2008) found that the short-spired species
of typical Conus (which would include C. hendersoni) all fall
into one clade (genus), Conus, whereas the taller-spired and, in many
cases, spirally sculptured species similar to the Recent species C. orbignyi
Audouin, 1831 and C. viminius Reeve, 1849 all fall into another clade
(genus), for which they used the genus Conasprella Thiele, 1929. The
relevance of this classification to the usage of the genus Conilithes
Swainson, 1840 for New Zealand tall-spired fossils deserves investigation, as
it is possible that Conilithes is simply an earlier name for Conasprella.
Meanwhile, the traditional name Conilithes is retained for
tall-spired fossils.
Distribution (3):
Tongaporutuan-Kapitean; Mangaotuwhito Stream, one km west of main
highway, northwest of Tikitiki, near East Cape, Kapitean (type); common and
well preserved in the shore platform at Maruhou Point, east of Te
Araroa, East Cape (figured), and poor specimens have been collected
from many localities in the Gisborne-East Cape district, mostly near East Cape.
Most records are from the shallow-water Tokomaru Sandstone facies, as
far south as Mangatuna
Quarry, inland from Tolaga Bay, and almost all records are Kapitean, but a few
are of Tongaporutuan
age (Late
Miocene).
One specimen probably assignable here is known from Upton Brook, Awatere
Valley (Kapitean).
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Conus
herdersoni
Maruhou Point, east of Te Araroa, East Cape,
Kapitean (M43165, National Museum) Late Miocene |
Conus herdersoni
mm. 60 Near Te Araroa, East Cape, North Island, New Zealand |
Conus
cf. kermadecensis
mm. 42 New Zealand (Femorale) |
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Conus
hendersoni
45mm long. From Miocene deposits, Maruhou Point, east of Te
Araroa, East Cape, North Island of New Zealand [ Mark
Large ] |
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Bibliografia Consultata